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Senior Applied Ai Engineer Startups Jobs

7,101 active opportunities · Updated for October 2026

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Pagerduty
📍 Lisbon• Full-time
1mo ago

PagerDuty (NYSE:PD) is a leader in Digital Operations Management. In an always-on world, organizations of all sizes trust PagerDuty to help them deliver a perfect digital experience to their customers, every time. Teams use PagerDuty to identify issues and opportunities in real time and bring together the right people to fix problems faster and prevent them in the future. Over 13,000 organizations (including 60 of Fortune 100) rely on PagerDuty to succeed with Digital Transformation, Cloud Migration, and DevOps Modernization. Notable customers include GE, Cisco, Genentech, Electronic Arts, Cox Automotive, Netflix, Shopify, Zoom, DoorDash, Lululemon and more. We are expanding rapidly as a platform for Digital Operations Management using AI/ML and Automation and growing our adoption by Development, IT, Customer Service, Security, and other teams across the organization. About the role PagerDuty’s Operations Cloud runs on a platform that ingests billions of signals and turns them into real-time action for thousands of customers. We’re looking for a Senior AI/ML Engineer who lives at the intersection of two disciplines: large-scale distributed systems and applied AI. In this role you will design and ship AI systems that run in production at PagerDuty’s scale — powering Incident Management AI Agents, event intelligence, and the LLM-powered capabilities embedded across our platform. You’ll own the full lifecycle, from framing the problem to serving reliably at scale. We are looking for a candidate who is genuinely passionate about building with modern AI — LLMs, agents, and retrieval — but grounded in the realities of building resilient, high-throughput systems. What you’ll do Design and build AI-powered features — LLM agents, retrieval, and event intelligence — that operate on high-volume, real-time event streams, from problem framing through production deployment and monitoring. Architect and own the systems behind them: agent and prompt orchestration, retrieval pipelin

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Coinbase
📍 - USA• Full-time• Remote• From $186.1K/yr
1mo ago

Ready to do the most impactful work of your career? At Coinbase , we are uncompromising on our mission to increase economic freedom. The bar is high, the environment is intense, and we like it that way. This isn't a place for complacency, it’s a place to be pushed past your perceived limits. If you're ready to build the future of finance alongside people who refuse to settle for "good enough," you belong here. Coinbase is a remote-first, but not remote-only company. Expect to get together quarterly for intense in-person working sessions called “surges.” learn more about working at Coinbase . As a Senior Software Engineer on the AI Platform team within the Platform group, you'll build and operate the LLM and agent infrastructure that every team at Coinbase depends on. This team owns the company's single path to large language models and the full agent lifecycle: build, deploy, run, observe, and improve. You'll lead multi-quarter technical initiatives across the platform, from gateway and runtime systems to knowledge bases and applied AI agents, directly shaping how Coinbase scales AI across the organization. What you'll do: Own the architecture and delivery of core platform systems including the LLM Gateway (60+ models, auth, PII redaction, fallbacks, cost optimization), AI Hub, and agent runtime with microVM sandboxes and governed MCP gateway Drive the design and implementation of Knowledge Base infrastructure, connecting data sources to auto-provisioned vector and markdown stores queryable by any agent Lead AI FinOps capabilities including spend attribution, governance, and cost optimization across all AI workloads company-wide Partner across engineering, security, legal, finance, product, and external partners at frontier labs and major cloud providers to ship high-impact platform capabilities Build evaluation and observability tooling including LLM-as-judge harnesses, full tracing, and feedback loops that let subject matter experts refine production a

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Scale AI
📍 San Francisco• Full-time• From $216K/yr
19 days ago

About Scale AI Scale AI is the data foundation for AI, helping organizations build and deploy reliable production AI applications. We partner with the world's leading enterprises and government organizations to accelerate their AI transformation through frontier AI systems that solve real business problems. Every day, we work with organizations across finance, healthcare, manufacturing, media and telecommunications to build production AI agents that automate complex workflows, help humans, reason over enterprise knowledge, and operate safely at scale. The Opportunity Applied AI is moving faster than ever. New foundation models, reasoning techniques, agent architectures, and research papers emerge every week. Yet building AI systems that reliably solve real-world problems remains one of the hardest engineering challenges. As a Senior Frontier Agent Engineer (Applied AI) , you'll bridge the gap between cutting-edge AI research and production deployment. You'll work directly with enterprise customers to design, evaluate, and deploy intelligent systems that combine frontier models with structured knowledge, retrieval, traditional machine learning, and enterprise software. Unlike traditional ML roles that focus on a single model or product, you'll work across a diverse portfolio of AI challenges spanning multiple industries and use cases. You may build a multi-agent research system and then participate in designing a customer intelligence platform, a healthcare copilot, or an autonomous workflow for a Fortune 100 company. If you enjoy reading new AI papers, experimenting with the latest models, and shipping production systems that create measurable business impact, you'll fit right in. What You'll Build Frontier AI Systems Design and deploy production AI agents that leverage the latest advances in large language models, reasoning, retrieval, memory, and tool use. Architect intelligent systems that combine LLMs, traditional machine learning, structured knowledge, enterpri

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Lyft
📍 Toronto• Full-time• From C$136K/yr
1mo ago

At Lyft, our purpose is to serve and connect. We aim to achieve this by cultivating a work environment where all team members belong and have the opportunity to thrive. Lyft is looking for software engineers from a scope of disciplines. We are growing our team with people who want to build, improve and incorporate technologies that make the lives of our community more enriched. As an engineer at Lyft, you'll collaborate with teams like product, data science, analytics, and operations on code that empower us to iterate quickly, while focusing on delighting our passengers and drivers. The Applied AI team is looking for a Backend Engineer to join the AI Entries team. You will build the foundational infrastructure that connects Lyft to emerging AI ecosystems and devices and architect the APIs and orchestration layers that allow third-party agents and multimodal interfaces to interact with Lyft. By building these robust integration, you will help make Lyft available where our riders are. Responsibilities: Establish engineering best practices and patterns; help uplift the team's craft and drive a culture of engineering excellence Drive high-impact projects and innovate new solutions to deliver the best user experience Produce and drive scalable system design for large, complex features — from idea through execution and launch Mentor engineers on the team, providing technical guidance and supporting their growth Write well-crafted, well-tested, readable, maintainable code Participate in code reviews to ensure code quality and share knowledge across the team Participate in the team's on-call rotation; identify, triage, debug, and resolve issues across our applications and platforms Have the ability to explain the various trade offs made in decisions Manage project priorities, deadlines, and deliverables. Experience: BS/MS or equivalent in Computer Engineering, Computer Science, or related field or equivalent practical experience. 5+ years of software engineering/production

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Synthesia
📍 London• Full-time• Remote
1mo ago

Synthesia is the world’s leading AI video platform for business, used by over 90% of the Fortune 100. Founded in 2017, the company is headquartered in London, with offices and teams across Europe and the US. As AI continues to shape the way we live and work, Synthesia develops products to enhance visual communication and enterprise skill development, helping people work better and stay at the center of successful organizations. Following our recent Series E funding round, where we raised $200 million, our valuation stands at $4 billion. Our total funding exceeds $530 million from premier investors including Accel, NVentures (Nvidia's VC arm), Kleiner Perkins, GV, and Evantic Capital, alongside the founders and operators of Stripe, Datadog, Miro, and Webflow. About the role As an Applied Research Engineer in our Video team, you will help build the next generation of production-grade foundation models for human-centric video generation. You will join a highly focused team working at the intersection of large-scale generative modeling, distributed systems, and production engineering. Our mission is to develop and optimize video base models that power realistic, controllable, and emotionally expressive synthetic humans at scale. This is not pure research. This is applied research with direct product impact. You will work on advancing training recipes, scaling distributed systems, improving evaluation frameworks, and optimizing inference to ensure our models are high quality, stable, and efficient enough for real-world deployment. Your work will directly influence models used by tens of thousands of businesses worldwide. What you’ll do You will own and execute end-to-end research and engineering projects, from hypothesis to production impact. This includes: Developing and scaling latent video diffusion models tailored for human-centric video generation Designing conditioning mechanisms to improve control (pose, emotion, script, camera) without sacrificing fidelity Advanc

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Synthesia is the world’s leading AI video platform for business, used by over 90% of the Fortune 100. Founded in 2017, the company is headquartered in London, with offices and teams across Europe and the US. As AI continues to shape the way we live and work, Synthesia develops products to enhance visual communication and enterprise skill development, helping people work better and stay at the center of successful organizations. Following our recent Series E funding round, where we raised $200 million, our valuation stands at $4 billion. Our total funding exceeds $530 million from premier investors including Accel, NVentures (Nvidia's VC arm), Kleiner Perkins, GV, and Evantic Capital, alongside the founders and operators of Stripe, Datadog, Miro, and Webflow. About the role As a Research Engineer in our Video team, you will help build the next generation of production-grade foundation models for human-centric video generation. You will join a highly focused team working at the intersection of large-scale generative modeling, distributed systems, and production engineering. Our mission is to develop and optimize video base models that power realistic, controllable, and emotionally expressive synthetic humans at scale. This is not pure research. This is applied research with direct product impact. You will work on advancing training recipes, scaling distributed systems, improving evaluation frameworks, and optimizing inference to ensure our models are high quality, stable, and efficient enough for real-world deployment. Your work will directly influence models used by tens of thousands of businesses worldwide. What you’ll do You will own and execute end-to-end research and engineering projects, from hypothesis to production impact. This includes: Developing and scaling latent video diffusion models tailored for human-centric video generation Designing conditioning mechanisms to improve control (pose, emotion, script, camera) without sacrificing fidelity Advancing distr

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1mo ago

Who we are At Twilio, we’re shaping the future of communications, all from the comfort of our homes. We deliver innovative solutions to hundreds of thousands of businesses and empower millions of developers worldwide to craft personalized customer experiences. Our dedication to remote-first work , and strong culture of connection and global inclusion means that no matter your location, you’re part of a vibrant team with diverse experiences making a global impact each day. As we continue to revolutionize how the world interacts, we’re acquiring new skills and experiences that make work feel truly rewarding. Your career at Twilio is in your hands. . Hiring and how we work We use Artificial Intelligence (AI) to help make our hiring process efficient. That said, every hiring decision is made by real Twilions! Also, while we are a remote-first company, you may be asked to report in person on an ad-hoc basis for team gatherings, functional off-sites or customer meetings. . See yourself at Twilio Join the team as Twilio's next Senior/Staff Applied Software Research Engineer for Emerging Technologies. We are hiring for multiple positions at a Senior and Staff level. About the job This full stack engineer position is needed to contribute hands-on technical expertise, collaborate on rapid prototyping, and develop innovative solutions as part of Twilio’s incubation team. As a Senior Applied Software Research Engineer, you’ll build hands-on skills with new technologies, collaborate with passionate engineers, and contribute to delivering solutions that help shape Twilio’s future offerings. This is a growth role, ideal for someone eager to learn and make a measurable difference in a fast-paced, entrepreneurial environment. As a Staff Applied Software Research Engineer, you’ll work closely with cross-functional partners to design and build new product ideas, leveraging modern technologies and your problem-solving abilities. This is a high-impact

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Asana
📍 New York• Full-time• $202K – $223K/yr
1mo ago

The AI Retrieval team powers the intelligence behind Asana's AI features by finding relevant work graph content and delivering it to LLM context windows. Our work enables AI features that truly understand your work—both within and outside of Asana—and use that understanding to take action. We also own Asana's traditional search experience. As a Senior Software Engineer on the AI Retrieval team, you'll build the systems that make Asana's AI smart and responsive. You'll tackle challenging problems in search and retrieval, working to improve the speed, cost-efficiency, and quality of our systems while expanding their capabilities to new data sources. This role is based in our New York City office with an office-centric hybrid schedule. The standard in-office days are Monday, Tuesday, and Thursday. Most Asanas have the option to work from home on Wednesdays. Working from home on Fridays depends on the type of work you do, and your recruiter can share more about the in-office requirements. What you'll achieve Reduce the latency and cost of our retrieval system, making Asana's AI features faster and more efficient Improve the quality and relevance of search results to help users find exactly what they need Expand the retrieval system's capabilities to query new Asana objects and third-party data sources Build and optimize search infrastructure using OpenSearch/ElasticSearch Contribute to ML-powered features like embeddings-based retrieval and semantic search Collaborate with cross-functional partners in New York City while building strong relationships with your Warsaw-based peers About you 6+ years of experience writing code in a production environment Curiosity about how AI and technology can solve problems and a desire to use it regularly in your work Hands-on experience in search engineering, including with OpenSearch or ElasticSearch Experience with embeddings and machine learning approaches to search and retrieval Experience in an Applied AI role is a plus Dem

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Linear
📍 United States• Full-time
1mo ago

At Linear, we're building the product development system for teams and agents. AI is fundamentally changing how software gets built, and we’re shaping the tools this new era requires. Founded in 2019, Linear has become the platform of choice for more than 40,000 companies (including OpenAI, Coinbase, and Ramp) to plan, build, and ship their products. Today, our team is distributed across North America, Europe, and Australia, and we’re continuing to grow internationally. What unites us is relentless focus, fast execution, and a deep care for software craftsmanship. We’re looking for experienced engineers who have shipped applied AI systems to production and want to define what the agent-native future looks like. We are building intelligence into the core of Linear, enabling the product to orchestrate coding, proactively move work forward, and power-up every software team. You’ll work closely with product and design to transform foundation models into structured, reliable workflows embedded deeply in the core of Linear. We care deeply about keeping Linear fast, intuitive, and opinionated—AI is no exception. Location & work mode Linear is a remote-first company, with optional co-working offices in San Francisco, New York, and London. This role is open to candidates based in the North America. You can work from anywhere within this region. We value deep focus and async collaboration, with intentional moments to connect in person through team off-sites, optional co-working, and occasional travel. What you'll do Build AI-powered product features that feel native, fast, and delightful to use Work with product and design to prototype and iterate on intelligent workflows and user interactions Design backend services to power natural language interfaces, smart suggestions, agentic workloads, and more Optimize prompts, fine-tune model behavior, and evaluate performance Help to guide our agent platform, allowing third parties to bring agents into the core Linear experience

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Sentry
📍 San Francisco• Full-time• $155K – $400K/yr
1mo ago

About Sentry Software runs the world and the pace is faster than ever. Sentry helps developers fix errors and performance issues before users notice, so teams can spend less time firefighting and more time building. Trusted by 200,000+ organizations, Sentry is today’s application monitoring standard and our team is building its AI-native future. About the role As a Senior Software Engineer on Sentry’s AI/ML team, you’ll be responsible for building the evaluation infrastructure that measures the accuracy, reliability, and real-world performance of our AI systems. This role is critical to ensuring that our debugging agents and AI-powered features behave correctly, safely, and predictably as they scale. You’ll design datasets, benchmarks, and test harnesses that turn ambiguous AI behavior into measurable signals, helping the team ship AI with confidence. In this role you will Design and build robust evaluation frameworks to measure accuracy, reliability, regressions, and edge cases in AI systems Create and curate high-quality datasets, golden test cases, and benchmarks grounded in real production data Build automated test harnesses and metrics pipelines to continuously evaluate models, prompts, and agentic workflows Partner closely with applied AI engineers and product leaders to define what “good” looks like and translate it into measurable criteria Own the evaluation lifecycle for major AI initiatives, from early experimentation through production monitoring You’ll love this job if you Care deeply about correctness, rigor, and measurement in AI systems Enjoy turning fuzzy product goals and model behavior into concrete tests and metrics Like building foundational infrastructure that unlocks faster iteration and higher confidence for the entire AI team Thrive in cross-functional environments and enjoy influencing model design through better evaluation Qualifications Minimum 5+ years of professional experience with a Bachelor’s degree in computer science, machine learni

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19 days ago

About Graphcore At Graphcore, we’re building the future of AI compute.We’re a team of semiconductor, software and AI experts, with deep experience in creating the complete AI compute stack - from silicon and software to infrastructure at datacenter scale.As part of the SoftBank Group, backed by significant long-term investment, we are delivering key technology into the fast-growing SoftBank AI ecosystem.To meet the vast and exciting AI opportunity, Graphcore is expanding its teams around the world.We are bringing together the brightest minds to solve the toughest problems, in a place where everyone has the opportunity to make an impact on the company, our products and the future of artificial intelligence. Job Summary As a Senior Machine Learning Engineer in the Applied AI team at Graphcore, you will contribute to advancing AI technology by developing and optimising AI models tailored to our specialised hardware. You will work on large scale systems where performance is critical to the success of our projects. Working closely with the Software development and Research teams, you will play a critical role in identifying opportunities to innovate and differentiate Graphcore’s technology. We seek engineers with strong technical skills and an understanding of AI model implementation at scale, eager to make a tangible impact in this rapidly evolving field. The Team The Applied AI team’s role is to be proxies for our customers, we need to understand the latest AI models, applications, and software to ensure that Graphcore’s technology works seamlessly with the AI ecosystem and at scale. We build reference applications, contribute to key software libraries e.g. optimising kernels for efficiency on our hardware, and collaborate with the Research team to develop and publish novel ideas in domains such as efficient compute, model scaling and distributed training and inference of AI models for multiple modalities and applications. If you're excited about advancing the next gen

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OpenAI
📍 San Francisco• Full-time
1mo ago

About the Team The Enterprise Identity team builds the identity foundation that enables organizations to adopt and use OpenAI products securely and reliably. The team owns the enterprise identity stack, including SSO, SCIM, tenant architecture, and identity capabilities across the enterprise admin experience and OpenAI's growing multi-product portfolio. About the Role We are looking for a hands-on senior technical leader to own the architecture and evolution of OpenAI's Enterprise Identity systems. You will set the long-term technical vision for the entire stack, establish shared identity primitives across products, and be accountable for systems that are foundational to our enterprise business. This role requires operating well beyond a single service or feature area. You will identify the most consequential architectural investments, align teams around durable solutions, and ensure our identity platform meets an exceptionally high bar for scale, availability, latency, and security. This role will be based in our San Francisco or Mountain View office. In this role, you will: Own the technical vision and architecture for the Enterprise Identity stack, including SSO, SCIM, tenant architecture, groups, permissions, and identity capabilities in enterprise administration surfaces. Lead the design and evolution of highly available, latency-sensitive identity systems serving a large and diverse global enterprise customer base. Establish common identity models and primitives that work consistently across OpenAI's products and enable the organization to scale. Set a high security bar by anticipating abuse cases, failure modes, and the long-term implications of new capabilities. Drive alignment across enterprise product, infrastructure, and security partners, resolving ambiguity and influencing roadmaps beyond the immediate team. Provide technical leadership to senior engineers and raise the quality of architecture and execution across the broader organization. You might thr

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OpenAI
📍 Washington• Full-time
1mo ago

About the Team Join the engineering teams that bring OpenAI’s ideas safely to the world! The Applied Engineering team works across research, engineering, product, and design to bring OpenAI’s technology to consumers and businesses. We seek to learn from deployment and distribute the benefits of AI, while ensuring that this powerful tool is used responsibly and safely. Safety is more important to us than unfettered growth. About the Role We’re seeking Software Engineers who can solve complex, high-impact problems across our stack. In this role, you’ll join a nimble team driving the deployment of OpenAI’s technology into new environments and infrastructure that power critical missions in the public sector. You’ll work cross-functionally with product, security, and compliance teams to build the functionality needed to deliver a scalable, reliable platform. You’ll also partner directly with customers to design and build new products and features that create real-world impact. From launching net-new capabilities to optimizing how we serve inference in unique, high-stakes environments, this role offers both breadth and technical depth—giving you the opportunity to shape the future of OpenAI’s technology where it matters most. This role is based in Washington D.C., San Francisco, CA or Seattle, WA. Occasional travel to customer sites is required for this role. In this role, you will: Own the development of new customer-facing ChatGPT and OpenAI API features end-to-end, both on-premises and in the cloud, for our public sector customers. Partner and directly embed with teams across the business, including engineering, security, and compliance, to enable our products to work within the unique constraints of new environments. Talk to users to understand their problems and design solutions to address them Work with the research team to get relevant feedback and iterate on their latest models, developing solutions specific for public sector customers at both the model & data

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Lyft
📍 Toronto• Full-time• From C$136K/yr
1mo ago

At Lyft, our purpose is to serve and connect. We aim to achieve this by cultivating a work environment where all team members belong and have the opportunity to thrive. Lyft is looking for software engineers from a scope of disciplines. We are growing our team with people who want to build, improve and incorporate technologies that make the lives of our community more enriched. As an engineer at Lyft, you'll collaborate with teams like product, data science, analytics, and operations on code that empowers us to iterate quickly, while focusing on delighting our passengers and drivers. The Applied AI team is looking for a senior software Engineer to join the Recommendations team. You will build the backend systems that decide what we surface to riders. You will architect the pipelines and services that turn rider context and signals into the right offer at the right moment, driving a measurable impact on rider experience. Responsibilities: Establish engineering best practices and patterns; help uplift the team's craft and drive a culture of engineering excellence Drive high-impact projects and innovate new solutions to deliver the best user experience Produce and drive scalable system design for large, complex features — from idea through execution and launch Mentor engineers on the team, providing technical guidance and supporting their growth Champion and evangelize the use of AI tools to accelerate engineering productivity across the team, sharing patterns and best practices that raise the bar for how the team builds Write well-crafted, well-tested, readable, maintainable code Participate in code reviews to ensure code quality and share knowledge across the team Participate in the team's on-call rotation; identify, triage, debug, and resolve issues across our applications and platforms Excellent communication skills and the ability to explain the various trade offs made in decisions Manage project priorities, deadlines, and deliverables. Experience: BS/MS or

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OpenAI
📍 San Francisco• Full-time
1mo ago

About the Team OpenAI’s Applications Engineering organization builds and operates the products that bring our cutting-edge research to millions of users and developers worldwide. The Applied Foundations team owns the core product and platform layers that make those experiences possible — from identity & access, to safety to payments & commerce across all of our apps. Our teams span product engineering, infrastructure, and safety, working together to deliver technology that is reliable, secure, and trusted at global scale. About the Role You will be a Senior Android engineer on OpenAI’s Applied Foundations team, building the core mobile experiences that power how users sign up, manage their account, family features, pay for services, stay safe, and interact with OpenAI’s products with confidence. This role is about creating high-quality products as well as reusable Android foundations that product teams across different OpenAI apps depend on to ship quickly while meeting the highest standards for security, reliability, and user trust. You’ll own complex client-side systems spanning UI, networking, local state, payment integrations and Apple platform integrations, and work closely with backend, product, and safety partners to shape the architecture that supports OpenAI’s mobile ecosystem at global scale. You might thrive in this role if you: Have 4+ years of professional software engineering experience. Have a proven track record of building high-quality Android applications in production. Are fluent in Kotlin (and/or Java) and familiar with Android development tools and architecture components. Prioritize performance, security, and user experience in mobile development. Enjoy working cross-functionally to bring ambitious product ideas to life. Care deeply about performance, security, and user experience. About OpenAI OpenAI is an AI research and deployment company dedicated to ensuring that general-purpose artificial intelligence benefits all of humanity. We

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